1959-1963 - Tustin Union High School, Tustin CA, diploma

1963-1967 - University of California, Santa Barbara, BA in Analytical Biology, with honors

1967-1971 - UCLA Medical School, PhD in PhysiologySupervisor: Jared M. Diamond

1971-1972 - Postdoctoral Fellow, Agricultural Research Council, Inst. of Animal Physiol., Babraham, Cambridge, UK Supervisor:  Richard D. Keynes  

1972 (3 mos) - Postdoctoral Fellow, II Physiologisches Institut der Universitat des Saarlandes, Homburg (Saar), Germany Supervisor:  Bernd Lindemann  

1973-1975 - Postdoctoral Fellow, Dept. Physiology-Anatomy Supervisor:  John G. Forte

1973-1975 - Lecturer, Dept. Physiology-Anatomy

1973 (3 mos) - Visiting Research Scientist. Departmento Biologia Cellular, Centro de Investigacion del Instituto Politecnico Nacional, Mexico, D.F.

1975 (3 mos) - Visiting Research Scientist, Physiological Laboratory, Cambridge, England

1975-1979  - Assistant Professor, Physiology-Anatomy Dept., University of California, Berkeley

1979-1985 - Associate Professor, Physiology-Anatomy Dept., University of California, Berkeley

1979 (3 mos) - Visiting Research Scientist. Panum Inst., Medical Physiol., Dept. A, Copenhagen, DK

1982-83 - Visiting Scientist (Sabbatical). Panum Inst., Med. Physiol., Dept. A, Copenhagen, DK

1985-90 - Key Investigator, Center for Ulcer Research and Education, UCLA and VA Wadsworth

1985-89 - Professor, Physiology-Anatomy Department, University of California, Berkeley

1989 - Professor of Cell and Developmental Biology, UCB

1990-93 - Head of Division, Cell and Developmental Biology, UCB

1967-71 - NIH Training Grant, predoctoral trainee Physiology Dept, UCLA

1971-73 - Bank of America-Giannini Foundation Postdoctoral Fellowship

1975 - Short Term Fellowship, European Molecular Biology Organization

2002 - Miller Institute for Basic Science Professor

AAAS

American Physiological Society

American Society of Cell Biologists

Biophysical Society

Society of General Physiologists

Editorial boards:

American Journal of Physiology, GI and Liver Physiology (1981 - 1997, 1999 - 2004)

American Journal of Physiology, Cell Physiology (1983 - present)

Associate Editor, American Journal of Physiology, GI and Liver Physiology, 1985 - 88

Ad hoc reviewer: American J. Physiology (Comparative Physiology Section and Lung Molecular and Cellular Biology Sections), Archives of Biochemistry, Biochemistry and Biophysics, Biochimica et Biophysica Acta, Biophysical J., Gastroenterology, Investigative Ophthalmology and Visual Science, J. Applied Physiology, J. Biological Chemistry, J. Cellular Physiology, J. Clinical Investigation, J. Comparative Physiology, J. Experimental Zoology, J. Membrane Biology, J. Neuroscience, J. Neurochemistry, J. Pharmacology and Experimental Therapeutics, Molecular Biology of the Cell, Nature, Pflugers Archiv, Physiological Reviews, Proceedings of the National Academy of Science, Proceedings of American Society of Experimental Biology and Medicine, Science

Merit Review Board for Basic Science (Study Section), Veteran's Administration, 1980-83

Ad hoc grant reviews for: NIH, NSF, Veterans Administration, Wellcome Trust (UK), Canadian Natural Sciences Research Council, Canadian Cystic Fibrosis Foundation, Cystic Fibrosis Foundation, Cystic Fibrosis Research Inc., Korean Natural Sciences

Ad hoc member NIH Study Sections: 1984, 1988, 1993, 2000, 2001, 2004, 2006, 2007

Pathobiology Study Section, Department of Defense Initiative in Breast Cancer, 1997, 1998, 2000

Secretary, Society of General Physiology, 1993-96

Councilor

American Physiological Society, GI section, 1990-92

Int'l Union of Physiological Sciences, 1990-92

Executive Board

CURE Digestive Diseases Center, UCLA, 1997-2000

Cystic Fibrosis Research, Inc, 1995-97, 2004-present